Flatten HTTP handler config (#2662)

Differentiating middleware and responders has one benefit, namely that
it's clear which module provides the response, but even then it's not
a great advantage. Linear handler config makes a little more sense,
giving greater flexibility and simplifying the core a bit, even though
it's slightly awkward that handlers which are responders may not use
the 'next' handler that is passed in at all.

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